Kokia cookei

O.Deg. (1934)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Malvales Family: Malvaceae Genus: Kokia

hau hele ‘ula [hau hele ula], koki‘o [kokio], kokia

Description

Key Characters:

Growth Form: Small trees perhaps 3–5 m tall.

Stems:

Roots:

Leaves: Leaves simple. Alternate. Blades 5–13 cm wide, 5–lobed or 7–lobed with shallow, angulate or deltate divisions. Base cordate to subcordate with an open sinus. Surfaces glabrate except lower surfaces tufted–pubescent in axils of major veins near base. Margins entire or nearly so. Petioles usually as Iong as or longer than blades, deciduous with prominent scars. Stipules caducous.

Flowers: Flowers solitary, clustered near the ends of the branches, axillary, pedicels stout, articulate near middle and sometimes with a persistent bract, terminal nectaries absent; pedicels 3–7 cm long, bracts up to 1.5 cm long, sometimes persistent; involucral bracts 3, distinct or connate basally, entire or lobed, broadly ovate, 25–35 mm long, 15–30 mm wide, shallowly to deeply 3–5 lobed, apex acute to obtuse, accrescent and leathery in fruit. Flowers bisexual (perfect). Calyx composed of connate sepals, cylindrical, 2-5-lobed, the lobes usually falling away leaving a basically truncate tube, tube ca. 1 cm long, the lobes valvate in bud. Corolla zygomorphic, corolla of 5 petals, orangish red, ca. 8 cm long and 6 cm wide, obovate, curved in bud, densely yellowish silky pubescent, reflexed above at anthesis, twisted at base, adnate at base to staminal column. Stamens 5 to numerous, monadelphous, forming a staminal column, staminal column ca. 6.5 cm long, exerted, curved; antheriferous in upper 1/3-1/2, terminated by 5 teeth; anthers monothecal. Pollen globose, echinate. Ovary superior, 5-celled; ovules 1 per cell, the carpels borne in a single whorl or rarely seemingly superposed whorls, placentation axile; style clavate, exceeding the staminal column, unbranched and stigmatically lobed at apex; stigma 5-lobed, sulcate, terminal or decurrent.

Fruit: Capsules depressed–globose to globose; up to 3 cm in diameter; woody; loculicidally dehiscent; apex apiculate. Seeds obovoid; 10–13 mm long; densely reddish lanate; with or without endosperm.

Ploidy: 2n = 24*

Habitat: Formerly endemic to the Mauna Loa area on the arid; western end of Molokai.

Elevation Range: ca. 450 m

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Only found in cultivation
O'ahu Only found in cultivation
Molokai Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Fl. Hawaiiensis 221: s.p. (1934)

Other References

Wagner et al. 1990:890 (Mo); Staples & Herbst 2005:383 (KEY), 393 (DESCR)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN j. f. c. rock 7076 NY 4/7/1910
2 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Judd, C.F. s.n. Molokai BISH 12/6/1923
3 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN [unknown] s.n. MEL
4 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN o. degener 8892 NY 3/7/1933
5 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information Cultivated. PRESERVED_SPECIMEN Liloa Dunn 161 Kauai PTBG 4/3/2003
6 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 3126 Maui PTBG 5/3/1994
7 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN o. degener, -. greenwell, -- hatheway & p. miller 21099 Maui US
8 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN L. M. Dunn 161 Kauai US 4/3/2003
9 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN l. m. dunn 161 Kauai US 4/3/2003
10 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN dunn, l. collector number: 161 Kauai US 4/3/2003
11 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN j. f. rock 7076 Molokai US
12 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J. F. 7076 Molokai US 4/10/2020
13 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Dunn, L. 161 Kauai US 4/3/2003
14 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Molokai US 8/8/1980
15 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Degener, O.; Greenwell, -.; Hatheway, --; Miller, P. 21099 Maui US 11/1/1950
16 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 27056 Molokai BISH 3/10/1955
17 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN US 8/7/1979
18 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wilder, G.P. s.n. Molokai BISH 4/15/1931
19 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information Cultivated. PRESERVED_SPECIMEN Tim Flynn 6809 Kauai PTBG 6/6/2001
20 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Judd Jr., A.F. 14 Molokai BISH 7/1/1925
21 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Cooke, G. 8982 Molokai BISH 3/1/1933
22 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Dunn, L. 161 Kauai BISH 4/3/2003
23 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Munro, G.C. 469 Molokai BISH 6/4/1916
24 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Cornelison, A.H. s.n. Oahu BISH 5/31/1945
25 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 7076 Molokai BISH 4/1/1910
26 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Dunn, L. Collector Number: 161 Kauai BISH 4/3/2003
27 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: s.n. Molokai BISH
28 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information W side of 0.4 ha islet, steep basalt wall. PRESERVED_SPECIMEN judd, c.s. collector number: 77 Molokai BISH 9/16/1937
29 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN wilson, s. collector number: sw 5 Oahu BISH
30 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 27056 Molokai BISH 3/10/1955
31 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN wilder, g.p. collector number: s.n. Molokai BISH 4/15/1931
32 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Conradt, C.C. s.n. Molokai BISH 7/1/1919
33 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Meyer s.n. Molokai BISH
34 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN cornelison, a.h. collector number: s.n. Oahu BISH 5/31/1945
35 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 5.mo Molokai BISH 6/1/1912
36 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 5.Mo Molokai BISH 6/1/1912
37 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN collector number: 124 Molokai BISH 6/27/1927
38 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN meyer collector number: s.n. Molokai BISH
39 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 7076 Molokai BISH 4/1/1910
40 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wilson, S. SW 5 Oahu BISH
41 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN conradt, c.c. collector number: s.n. Molokai BISH 7/1/1919
42 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN cooke, g. cooke, mrs.g. collector number: 8982 Molokai BISH 3/1/1933
43 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN judd jr., a.f. collector number: 14 Molokai BISH 7/1/1925
44 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN judd, c.f. collector number: s.n. Molokai BISH 12/6/1923
45 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN lyon, h.l. collector number: s.n. Molokai BISH 4/12/1929
46 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information Dry regions. PRESERVED_SPECIMEN judd, c.s. collector number: 77 Molokai BISH 9/16/1937
47 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lyon, H.L. s.n. Molokai BISH 4/12/1929
48 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information Grassy, shrubby ridge PRESERVED_SPECIMEN j. f. c. rock s.n. NY 3/7/1910
49 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information Dry regions. PRESERVED_SPECIMEN Judd, C.S. 77 Molokai BISH 9/16/1937
50 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN munro, g.c. collector number: 469 Molokai BISH 6/4/1916
51 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN O Degener|A Greenwell|W H Hatheway|H A Miller AK
52 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN alwyn h. gentry|rosa ortiz gentry 78013 MO 8/12/1992
53 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN waimea arboretum waimea arboretum s.n. #18 MO 9/15/1997
54 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN o. degener 21099 NY 11/7/1950
55 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN d. w. orr s.n. NY 8/7/1990
56 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN otto degener and isa legmer LSU 11/8/1959
57 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information Cultivated. PRESERVED_SPECIMEN Hank Oppenheimer H81418 Maui PTBG 8/28/2014
58 Kokia cookei O.Deg. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN 124 Molokai BISH 6/27/1927